The year is 1868; English nobleman George Joestar and his son Jonathan become indebted to Dario Brando after being rescued from a carriage incident. What the Joestars don't realize, however, is that Dario had no intention of helping them; he believed they were dead and was trying to ransack their belongings. After Dario's death 12 years later, George—hoping to repay his debt—adopts his son, Dio. While he publicly fawns over his new father, Dio secretly plans to steal the Joestar fortune. His first step is to create a divide between George and Jonathan. By constantly outdoing his foster brother, Dio firmly makes his place in the Joestar family. But when Dio pushes Jonathan too far, Jonathan defeats him in a brawl. Years later, the two appear to be close friends to the outside world. But trouble brews again when George falls ill, as Jonathan suspects that Dio is somehow behind the incident—and it appears he has more tricks up his sleeve.

After graduating from a top university with an impressive extracurricular record in the rugby club, Akira Tendou has nailed every step of the way to securing his dream job. On top of that, a beautiful and kind co-worker always brightens his day in the office! Life seems to be going very well for Akira until he slowly realizes that sleepless nights and brutal work are his new reality. Due to three years of mind-numbing labor in an exploitative company, Akira is unable to recognize the tired, unaccomplished person he has become. On track to losing all passion in life like several of his overworked colleagues, Akira finds his saving grace in the most unexpected way possible—the breakout of a zombie apocalypse. With the free time he finally has, Akira decides to complete a bucket list of a hundred things he wants to do before he eventually gets turned into a zombie. Although he is surrounded by the dead, Akira has never felt more alive!

Rugby is a sport where anybody can be a star when they hold the ball. The rules are simple: one player has to carry the ball past the goalposts across the field while avoiding the other team, who are allowed to use all their force to knock the enemy down. However, the player can pass to a teammate behind them or kick the ball before they are tackled. Unlike other sports, the game continues even when the person holding the ball is taken down. Kenji Gion is a short but feisty high school freshman at Kanagawa High School. Right after the school's opening ceremony, he meets the tall and timid Sumiaki Iwashimizu, and the two of them get an invitation to observe the rugby club. When Gion watches the match, he realizes that even though big and tall guys have an advantage, there are positions for people of all shapes and sizes. Immediately convinced that he has found a sport he can excel at despite his short stature, he signs up, though Iwashimizu declines. Gion is unable to accept Iwashimizu's excuses and drags him back into the sport. Their training becomes intense as they aim to become the champions of Hanazono, the national high school rugby championships.

Natsuza Yuzuki has been playing rugby as a left-winger ever since he was a child. After enrolling in Doushisha University, he quickly becomes a rising star in the Kansai University League. His dreams are shattered, however, when a nasty motorbike accident leaves him with a spinal hernia—making him unable to be able to play rugby ever again. With no role available other than the position of manager, he accepts his new post and hopes his squad can obtain a league title. Yuzuki's team is an unlikely group, including his stoic best friend and full-back Seiichirou Jingyouji; hot-headed first-year Yasunari Tsuru, who harbors a strong dislike for Natsuza, filling his senior's old position as left wing; and Yuu Mashiro, who is struggling to hold his ground as a scrum half. Despite Natsuza putting his all into training his team for a league title, their squad still feels incomplete. A year ago, Ibuki Ueka, the genius fly-half that was once Doushisha's ace, suddenly quit rugby—and now Natsuza is determined to get him back.