Sakura and Syaoran live peaceful lives in the desert country of Clow. Despite the King's disapproval of the budding romance between the princess and her archeologist friend, their days remain blissful if not a little dull. But that soon changes when Sakura appears in front of Syaoran in a trance with wings on her back. When the wings dissolve into feathers and Sakura loses consciousness, Syaoran is spurred to action. Syaoran soon learns that he must collect Sakura's feathers, which have been scattered across many different realms, in order to protect her life. After making a deal with the Dimensional Witch Yuuko Ichihara, Syaoran teams up with Mokona Modoki, a creature who has the power to traverse realms. Also accompanying him on this journey are Fai D. Flourite, a carefree man fleeing persecution in his home world, and Kurogane, a swordsman forced to travel to refine his strength and character. Together, the bizarre alliance works to reclaim Sakura's feathers—no matter how impossible it seems to find them all.

With a new school year comes a new crowd of classmates, and for their final year of junior high, aspiring writer Kotarou Azumi and track team member Akane Mizuno end up in the same class. Though initially complete strangers, a few chance encounters stir an innocent desire within their hearts. A yearning gaze, a fluttering heart—the hallmarks of young love slip into their lives as fate brings their paths to a cross. However, though love is patient and love is kind, Kotarou and Akane discover it is not always straightforward. Despite the comfort they find in each other's company, heartache and anxiety come hand in hand with pursuing the feelings in their hearts. With the uncertainty of how the other truly feels as well as the competing affections of those around them, the road ahead is unclear. Even so, under the shining light of a beautiful full moon, Kotarou gathers his courage to ask Akane a single question, one that forever changes their quiet relationship.

In Japanese folklore, a tsukumogami is an object that has gained a soul, becoming alive and self-aware. There are two types of tsukumogami: the mature tsugumomo, who have developed through long years of harmony with their owners, and the aberrant amasogi, premature spirits that are only born to grant the destructive wishes of certain people. Kazuya Kagami has never gone without his mother's obi after her death. Be it at home or school, he keeps it safe with him at all times. One day, he nearly loses his life when a wig amasogi attacks him. When all seems to be over, his treasured obi defends him, transforming into a beautiful girl. She introduces herself as Kiriha, a tsugumomo owned by Kazuya's mother. With Kiriha's arrival, Kazuya enters a reality he has never seen before, a world with gods and tsukumogami.

A series of unaired comedic shorts released with Blu-ray and DVD boxes of Tsuki ga Kirei.