Another installment in the Tengen Toppa Gurren Lagann Parallel Works anthology series of animated shorts, featuring characters from Gurren Lagann in alternative worlds. Every clip is animated by a particular animator of GAINAX. Creators are Gekidan Inu Curry, Akemi Hayashi, Sushio, Shouko Nishigaki, Satoru Yamaguchi, Shouji Saeki & Shingo Abe and Ayumu Kotake.

Humans have enjoyed their lavish, peaceful, and prosperous lives for seven years since the day the almighty Spiral King was defeated—the day they reclaimed their homeland, Earth. However, the boon of this lifestyle leaves them unprepared when an unknown, hostile threat arises due to the ever-growing human population. This calamity is the Anti-Spiral—a fearsome enemy with unparalleled power. As the Spiral King's prognosis postulating the destruction of The Spiral's World begins to come true, the pieces are in place, and Team Dai-Gurren is ready. With his late brother's hope to see a better future for mankind, Simon—along with Nia Teppelin and the rest of the team—is determined to overthrow the mighty Anti-Spiral in order to revive humanity's lost hope.

Simon and Kamina were born and raised in a deep, underground village, hidden from the fabled surface. Kamina is a free-spirited loose cannon bent on making a name for himself, while Simon is a timid young boy with no real aspirations. One day while excavating the earth, Simon stumbles upon a mysterious object that turns out to be the ignition key to an ancient artifact of war, which the duo dubs Lagann. Using their new weapon, Simon and Kamina fend off a surprise attack from the surface with the help of Yoko Littner, a hot-blooded redhead wielding a massive gun who wanders the world above. In the aftermath of the battle, the sky is now in plain view, prompting Simon and Kamina to set off on a journey alongside Yoko to explore the wastelands of the surface. Soon, they join the fight against the Beastmen, humanoid creatures that terrorize the remnants of humanity in powerful robots called Gunmen. Although they face some challenges and setbacks, the trio bravely fights these new enemies alongside other survivors to reclaim the surface, while slowly unraveling a galaxy-sized mystery.

Ever since his powers awakened, Tenchi Masaki has attracted the attention of many powerful galactic beings, including the mysterious Lady Takami who dispatches her best warrior, Z, to observe him. Though he is under strict orders not to harm Tenchi, Z has no intention of playing by the rules and begins plotting his demise. Meanwhile, on Earth, Tenchi tries to piece together more information about his family and the universe. But as usual, his life won't be getting any easier, as his entire household enters a state of panic when Noike Kamiki Jurai—a woman claiming to be Tenchi’s fiancée—shows up out of the blue.