Due to her father's remarriage, robust 16-year-old Himeno Awayuki moves into a large mansion with a beautiful garden—the quintessential dream house for any girl her age. However, much to Himeno's disappointment, her new stepfamily doesn't really seem to like her, as her stepmother often occupies herself with her father, her younger stepsister Mawata ignores her, and her other stepsister—the equally aged Mayune—tries to prank her at every opportunity. But Himeno doesn't have time to dwell into thoughts of hopelessness—her new life has now become involved with a group of seven magical boys known as the Leafe Knights, after they ask her to become a magical princess who can borrow their powers! Although Himeno accepts their request and becomes the Prétear, she feels doubtful in her abilities to protect the world and its Leafe, the source of energy for all life. Will Himeno be able to find happiness among her new family and also save the Earth from the enemy, the Princess of Disaster?

Ryouma Echizen is one of 50 nationally ranked middle school tennis players to receive an invitation for the Japan U-17 tennis training camp. Previously open only to high school players, the exclusive regiment trains the best players in the country for the upcoming U-17 World Cup. The high schoolers are not pleased that these middle schoolers are allowed into the camp, but the middle schoolers easily overwhelm them before they are halted by the coaches. The rules of the camp are soon explained: the players are split into one of 16 courts based on skill, with the best players occupying Court 1. Players move up and down courts based on shuffle matches that occur before practice begins each day. In anticipation of a game, the middle schoolers are asked to pair up with one another, only to find out that they would not be playing doubles with their partners. Instead, they are pitted against each other in a tiebreaker-style game. The winner of the match would be allowed to stay to further develop their skills whereas the loser would be sent home. For the sake of tennis, friendships and camaraderie are all put on the line. As the fierce competition between the middle and high schoolers persists, the situations they find themselves in goes deeper than just playing to remain in the camp.

The Atobe Group is establishing a new tennis court in Kanagawa and is planning to celebrate its opening with an exhibition match. Hyoutei Academy's tennis team captain, Keigo Atobe, nominates the best team in the region to participate: Rikkai University-Affiliated Middle School. On behalf of the Rikkai tennis team, Seiichi Yukimura accepts the challenge to showcase the prowess of its members. Amidst tense preparations, the third-years of Hyoutei supervise next year's captain, Wakashi Hiyoshi. At the same time, the third-year members of Rikkai begin to worry whether Akaya Kirihara, the team's lone second-year member, can lead the club to glory once again. Determined to overthrow their respective captains' legacies during the match, both Hiyoshi and Kirihara continue rigorous training to prove they have what it takes to succeed them. The future of Hyoutei and Rikkai rests on the shoulders of these second-year aces, who harbor high expectations and ambitious goals. Emboldened by their endeavors, the third-years of each respective team impart their feelings to the second-years that will be carrying the torch for their schools once they graduate.

With the arrival of the U-17 World Cup, youth tennis teams from all around the world gather in Melbourne, Australia to determine which is the best. With nine-time champion Germany vying to defend their title once more, and powerhouse nations—Switzerland, France, and the United States—determined to make things interesting, the road to victory is perilous. Although Japan is sitting at the bottom of the rankings, the addition of the nation's strongest middle school players has reinvigorated their chances at winning. Captained by the ruthless Houou Byoudouin, Japan is ready to show the world what they are capable of. However, before the real competition starts, the teams will have a chance to prove their strength through exhibition matches. As luck would have it, Japan is slated to face off against defending champion Germany—a match they are likely to lose. To make matters worse, old allies have suddenly reappeared as sworn enemies. Before Team Japan crumbles under the pressure of the international stage, they must rise from the ashes and claim the title for themselves.