It is A.C. 196—one year after the conflict between Earth and its nearby space colonies ended with the formation of the Earth Sphere Unified Nation. As mankind enters a new era of unity and prosperity, four of the five Gundam pilots decide to follow in society's footsteps and disarm by launching their mobile suits into the sun. However, their plan is unexpectedly suspended when a rebellion erupts with the emergence of Mariemaia Khushrenada—who claims to be the child of fallen dictator Treize and desires to conquer the world in his stead. Deciding that they are the last line of defense for the Earth Sphere, Heero Yuy, Trowa Barton, Duo Maxwell, and Quatre Raberba Winner scramble to retrieve their Gundams and fight to protect the peace the people of the colonies have pursued for so long. Despite their unwavering resolve to complete one final mission, the battle will be challenging: the pilots' former ally Wufei Chang has pledged his allegiance to Mariemaia's cause instead.

It has been 33 years since an alien race, known as the JAM, emerged from a hyperspace passage over Antarctica with the sole purpose of invading Earth. Despite their unexpected initial assaults, a combined effort by humanity not only pushed the JAM force off of Earth but also moved the frontlines of the battle onto the JAM's homeworld of Fairy. Though out of sight and mind for the majority of Earth's inhabitants, the conflict still rages on Fairy, where the United Nations has established five bases and a powerful, technologically advanced military—including the Fairy Air Force (FAF). Lieutenant Rei Fukai acts as a pilot for the FAF Special Air Force’s Boomerang reconnaissance squadron, flying a Super Sylph fighter jet with the personal name Yukikaze. Soaring high above and observing the battles for air superiority, he is responsible for collecting and analyzing valuable intelligence on JAM methods. While his flying skills are unmatched, he also heavily relies on the highly advanced weaponry and near-sentient AI system of his aircraft. Though despised by other members of the FAF for his antisocial and standoffish nature, Lt. Fukai continues to carry out his duties along with Yukikaze. However, during a particularly bloody battle, the appearance of an unresponsive and hostile aircraft catches the lieutenant off-guard—an aircraft which is of the same make and model as his own.

Average high school student Kyou Sogoru is an avid swimmer living in beautiful Maihama City. He spends his days hanging out with friends, swimming, and playing video games. However, his normal life turns upside down when a beautiful and mysterious girl named Shizuno Misaki approaches him with a strange request—jump into their school's pool together. This fateful leap transports Kyou into a war-torn world where humans pilot impressive humanoid robots known as Zegas to fight against malicious aliens known as Gards-Orm. To spearhead this endeavor, humans have formed Cerebrum, a rebel organization working to prevent the Gards-Orm from eradicating humankind. As Kyou participates in combat operations, meets his fellow Zega pilots, and witnesses countless deaths, he begins to question the true nature of this world as well as his own life.

What happens when two people love each other but just aren't meant to be together? Takaki Toono and Akari Shinohara are childhood friends, but circumstances beyond their control tear them apart. They promise to stay in contact, and although the progression of time widens the distance between them, the chain of memories remains ever-present. Byousoku 5 Centimeter is a romantic drama that focuses on the mundane and harsh reality of long-distance relationships. Stuck in the past and unable to make any new memories, Takaki and Akari cling to the hope of seeing each other again. They live their everyday lives half-heartedly, both hurting themselves and the people around them.