The Mikadono sisters—Miwa, Niko, and Kazuki—stand at the pinnacle of Saika Academy as its Three Royals, renowned respectively as prodigies in shogi, karate, and theater. Unfortunately, they have drifted apart from each other due to their overwhelming dedication to their fields, effectively turning them into strangers living under the same roof. Enter Yuu Ayase, the son of the late Subaru Ayase, a legendary actress who graduated from Saika. Unlike his mother, Yuu lacks any extraordinary talent whatsoever, much to the disappointment of those who knew her. Still, he transfers to her alma mater with one quiet promise in his heart: to fulfill her final wish that he build a true and happy family. It just so happens that the Mikadono patriarch is also Subaru's longtime friend and benefactor, and he offers Yuu both a place to stay and a fresh start. Thrust into the lives of three estranged sisters who act cold toward him, Yuu may not be a genius himself, but the clumsy sincerity he brings might be what they all need to start anew.

Since childhood, Hana Haruyama has been enthralled by the art of storytelling. After watching a child her age perform on television with mesmerizing grace, Hana decides to try it herself. She practices by reciting tales to Tonaki Island's children, who are always enthusiastic about her performances. However, growing up in such a small community, Hana remains unaware of the possibilities beyond her world, satisfied with how things are. Entering Sumomogaoka High School as a first-year student, Hana catches the attention of Mizuki Usurai, the president of the broadcasting club, with her skills. Despite Hana's hesitation and anxiety about the road ahead, Mizuki successfully convinces her to join the club, nurture her gift, and perhaps one day reach the top—the nationals of the NHK Broadcasting Competition.

After a disastrous romantic confession, Masamune Okumura finds solace in the fictional world of anime and manga. Now a second-year high school student and the president of the Manga Research Club, Masamune spends a peaceful existence watching the adventures of the angel Liliel, his favorite character. In the beginning of the academic year, Masamune’s life is turned upside down when he meets first-year student Ririsa Amano, a passionate cosplay practitioner. Despite Masamune's initial reluctance, the determined young woman manages to convince him to become her personal photographer for the production of a collection dedicated to their common favorite character: Liliel. As Masamune enthusiastically discovers the world of cosplay and photo editing, he is unexpectedly reunited with his childhood friend Mikari Tachibana, who has become a model in the hope of impressing her beloved upperclassman. Realizing that modeling is not enough to intrigue the hopeless otaku, Mikari decides to join the club and become a cosplayer herself, all to finally have a chance at winning Masamune's heart. However, Masamune's love for fictional characters may not be capable of extending to those who impersonate them.

While helping out at an anime convention to promote his city, high school student Kakeru Niimi is caught up in a sudden earthquake. During the emergency, he ends up entering a shrine where he sees a peculiar relic shatter. The very next day, his classmate Miyako Kujou, who was cosplaying at the event, worries that she has been cursed as a result of the earthquake, but Niimi brushes her suspicions off. However, Niimi's mind changes when he discovers that various people have gained supernatural powers in the city—and that some have started using them for nefarious purposes. Fortunately, Niimi is not alone: he is joined by Miyako, his sister Sora, and a cold girl named Noa Yuuki, who have all awakened to their own abilities. If the group wants to keep their hometown safe, they must strive to prevent any future incidents while unraveling the mystery behind it all.