Twintails: the glory of all mankind—or at least that is what first-year high school student Souji Mitsuka believes. At school, Souji spends most of his time daydreaming and rating girls' twintails, even going as far as creating a club dedicated to the hairstyle. His obsession does not go unnoticed, however; when monsters from outer space attack Earth and claim the world's twintails for themselves, a strange woman named Twoearle enlists Souji to fight back using twintails of his own! By transforming into the twin-tailed warrior Tail Red, Souji combats the vicious alien organization known as Ultimegil, whose main goal is to colonize Earth and steal everyone's spiritual energy, or attribute power. Alongside fellow twin-tailed fighters Aika Tsube and Erina Shindou, Souji must find a way to defeat the invading army and defend the twintails he holds so dear.

Mahiru Shiina is worthy of her nickname Angel: she is a divine beauty loved by all, and she excels in both academics and athletics. Shiina lives in an entirely different world from Amane Fujimiya, her next-door neighbor. Despite living so close together, they have never spoken once. But their silence is broken when Fujimiya spots Shiina gloomily sitting on a swing amidst a heavy rainstorm and lends her his umbrella. When Fujimiya catches a cold the next day, Shiina wishes to return the favor for the umbrella by nursing him back to health. Believing that this would be their first and last interaction, he silently appreciates her kindness. However, Shiina—who cannot help but worry about Fujimiya's lack of tidiness and proper nutrition—begins to cook and clean for him. As the unlikely pair spend time together in Fujimiya's apartment, they explore the true nature of their relationship and the gentle emotions that emerge from it.

Mikoto Shinozaki is a high school student lacking in social awareness. In addition, due to his untidy appearance and low self-esteem, he is an easy target for bullies. Regardless, any misfortune that falls upon Mikoto is partly his own fault—at least, that is what Yuki Fukuzawa, the most popular girl in class, believes. With this assumption, Yuki invites Mikoto to join the school's bicycle club, in hopes he can improve himself and gain new friends. Motivated by feelings he has for Yuki, Mikoto has no reason to decline her offer—except for the fact that he does not know how to ride a bicycle. Despite his lack of even the fundamentals, Mikoto cannot get over the image of himself on a bicycle. Eventually, he succumbs to his curiosity and sneaks into the school grounds at night to practice. This begins Mikoto's journey as he pedals his way through the vast world of cycling, where the scenery of his life changes in ways he never thought possible.

Shinichi Kanou is a shut-in otaku with a vast knowledge of anime, manga, and video games. One day, after applying for a job in hopes of escaping his secluded lifestyle, he is kidnapped and transported to the Eldant Empire—a fantasy world filled with elves, dragons, and dwarves. Trapped in this strange land, Shinichi is given an unlikely task by the Japanese government: to spread otaku culture across the realm by becoming an Otaku Missionary. To accomplish his mission, Shinichi has the full support of the Japanese government, as well as the half-elf maid Myucel and Princess Petralka of the Eldant Empire. Together with this ragtag bunch, he will overcome the obstacles of politics, social classes, and ethnic discrimination to promote the ways of the otaku in this holy land.