The homeland of the Genesect exists no more. After three hundred million years, the snowy mountains where the Paleozoic Pokémon once lived are no longer habitable—but not all of the race are actually extinct. Several specimens are still alive and desperate to find a new place to call home. Led by a Shiny Genesect and accompanied by the Legendary Pokémon Mewtwo, the vagabond creatures arrive in New Tork City. Perceiving the city skyline as similar to their initial habitat, the Genesect take over the city center, driving out other Pokémon and threatening the existence of the local populace. On top of that, the Genesect leader confronts Mewtwo in a cunning, fast-paced duel that is bound to destroy the entire city—and it's up to Satoshi and his companions to save the city and reconcile the feuding Pokémon.

In this latest cinematic adventure, Satoshi meets the Mythical Pokémon Volcanion when it crashes down from the sky, creating a cloud of dust—and a mysterious force binds the two of them together! Volcanion despises humans and tries to get away, but it’s forced to drag Satoshi along as it continues its rescue mission. They arrive in a city of cogs and gears, where a corrupt minister has stolen the ultimate invention: the Artificial Pokémon Magearna, created 500 years ago. He plans to use its mysterious power to take control of this mechanical kingdom! Can Satoshi and Volcanion work together to rescue Magearna? One of the greatest battles in Pokémon history is about to unfold! (Source: Official site)

While visiting a big city home to many lively Pokémon, Pikachu rejoices when Satoshi, Kasumi, and Takeshi give their Pokémon free rein to play, with the only condition that they must all be at this rooftop in two hours. Unfortunately, Pikachu's leisure time is short-lived when he encounters a Pichu on a rooftop across the street. While bantering with each other, Pikachu nearly falls off the building. Following a lucky sequence of events, Pikachu finds himself safe next to Pichu and his brother—but on the opposite side of the road! Because they are familiar with the area, the Pichu brothers volunteer to escort Pikachu back to his friends. As the trio stumble upon a series of obstacles on their journey across the bustling city, Pikachu starts to wonder if he will rejoin his companions on time.

After his mother wins a free trip to the islands, Pokémon trainer Satoshi and his partner Pikachu head for Melemele Island of the beautiful Alola region, which is filled with lots of new Pokémon and even variations of familiar faces. Eager to explore the island, Satoshi and Pikachu run wild with excitement, quickly losing their way while chasing after a Pokémon. The pair eventually stumbles upon the Pokémon School, an institution where students come to learn more about these fascinating creatures. At the school, when he and one of the students—the no-nonsense Kaki—have a run-in with the nefarious thugs of Team Skull, Satoshi discovers the overwhelming might of the Z-Moves, powerful attacks originating from the Alola region that require the trainer and Pokémon to be in sync. Later that night, he and Pikachu have an encounter with the guardian deity Pokémon of Melemele Island, the mysterious Kapu Kokeko. The Pokémon of legend bestows upon them a Z-Ring, a necessary tool in using the Z-Moves. Dazzled by their earlier battle and now in possession of a Z-Ring, Satoshi and Pikachu decide to stay behind in the Alola Region to learn and master the strength of these powerful new attacks. Enrolling in the Pokémon School, Satoshi is joined by classmates such as Lillie, who loves Pokémon but cannot bring herself to touch them, Kaki, and many others. Between attending classes, fending off the pesky Team Rocket—who themselves have arrived in Alola to pave the way for their organization's future plans—and taking on the Island Challenge that is necessary to master the Z-Moves, Satoshi and Pikachu are in for an exciting new adventure.