Grief-stricken over the death of his last family member, his dog Hanako, picture book writer Hotaru Yuuyami suffers from writer's block for nearly two months. Despite his childhood friend and editor Tsubaki Uehara's best efforts, nothing seems to alleviate his pain. But all of this changes one day with the sudden apparition of Kinoko Inu, a mysterious mushroom from his garden that turns into a dog-shaped animal. Much to the great dismay of Tsubaki, Hotaru hastily invites the creature into his abode. Days after Hotaru and Kinoko Inu start living together, the strange mushroom-dog decides to burn the last of Hanako's personal belongings in order to alleviate Hotaru's depression. However, the cohabitation between the overworked writer and his mischievous guest proves more difficult than expected, and Hotaru must let go of his grief if he wants to properly take care of Kinoko Inu.

While the perfect getaway for most girls her age might be a fancy vacation with their loved ones, Rin Shima's ideal way of spending her days off is camping alone at the base of Mount Fuji. From pitching her tent to gathering firewood, she has always done everything by herself, and has no plans of leaving her little solitary world. However, what starts off as one of Rin's usual camping sessions somehow ends up as a surprise get-together for two when the lost Nadeshiko Kagamihara is forced to take refuge at her campsite. Originally intending to see the picturesque view of Mount Fuji for herself, Nadeshiko's plans are disrupted when she ends up falling asleep partway to her destination. Alone and with no other choice, she seeks help from the only other person nearby. Despite their hasty introductions, the two girls nevertheless enjoy the chilly night together, eating ramen and conversing while the campfire keeps them warm. And even after Nadeshiko's sister finally picks her up later that night, both girls silently ponder the possibility of another camping trip together.

Mystery Camp Relaxing in the heart of nature is a constant joy for Nadeshiko Kagamihara, Aoi Inuyama, and their friends. But when the camping experience changes in unexpected ways, one can only be curious about the extraordinary wonders that have yet to be explored. Tabisuru Shima Rin Gathered around a campfire, the girls of the Outdoor Activities Club and Ena Saitou all propose various ideas to Rin Shima for her next trip. However, their suggestions are tailored to their personal preferences—not to mention that Rin has plans of her own.

Having spent Christmas camping with her new friends, Rin Shima embarks on a solo-camping trip to see the New Year sunrise by the sea. All goes according to plan until unforeseen weather blocks the roads back home, making a return trip impossible. Rin, who is now stranded for a few days, is invited by Nadeshiko Kagamihara to stay at her grandmother's house. What is supposed to be a two-day trip becomes an extended period of sightseeing and new experiences for Rin, and she encounters some new and old faces along the way. Yuru Camp△ Season 2 continues the story of Rin, Nadeshiko, and their friends as they further explore the joys of camping.