Having spent Christmas camping with her new friends, Rin Shima embarks on a solo-camping trip to see the New Year sunrise by the sea. All goes according to plan until unforeseen weather blocks the roads back home, making a return trip impossible. Rin, who is now stranded for a few days, is invited by Nadeshiko Kagamihara to stay at her grandmother's house. What is supposed to be a two-day trip becomes an extended period of sightseeing and new experiences for Rin, and she encounters some new and old faces along the way. Yuru Camp△ Season 2 continues the story of Rin, Nadeshiko, and their friends as they further explore the joys of camping.

Yurume, an 18-year-old ronin and recent high school graduate, has high hopes of being accepted and attending Tokyo University. She moves into an apartment complex, the Maison du Wish, located on the outskirts of Tokyo where other ronins living there are studying for upcoming college acceptance exams.

As time marches forward, Rin Shima, Nadeshiko Kagamihara, and their friends have grown into full-fledged adults. Now with careers and responsibilities, they can no longer freely spend their days camping and adventuring. However, Chiaki Oogaki, who currently works for the Yamanashi Tourism Organization, reunites the group as she comes up with an exciting proposal—to build a campground together to revitalize an abandoned patch of land from their homeplace, the Yamanashi Prefecture. Motivated by this ambitious project, the girls combine their efforts to create an environment where future generations can enjoy unique moments as they once did, all while reminiscing the charms of camping from their high school days.

In the cyber city Tom Sawyer, society revolves around earning Love, akin to receiving likes on social media. Citizens of this augmented reality exchange Love for public services and appearance-changing avatars via the Decoration Customizer, also known as Deco, a hologram device implanted in an individual's eyes from early childhood. Under the government's careful management, the people of Tom Sawyer live a peaceful existence, outside the occasional appearance of Phantom Zero, a mysterious criminal who brings all Love to zero within their vicinity. Berry, a mischievous girl fascinated by Phantom Zero, lives comfortably in the system of Tom Sawyer until her Deco malfunctions, enabling her to see a camouflaged prankster named Hack. Believing Hack to be Phantom Zero, Berry chases after the suspect and, in the process, begins to uncover society's most well-kept secrets. As it turns out, Hack is not the only person living outside the system's rules—and the vibrant colors of Tom Sawyer hide a darker nature.