A great evil is sweeping over the realm; an evil that the young swordsman Roan and his life-long companion, the acolyte Yufa, must face head on! For these two travel toward their destiny, from the highest towers to the depths of the underworld, through forest and desert alike. With an ever-growing cast of fellow heroes, fate will grasp these travelers by their very souls and propel the band of skilled adventurers towards a noble end. Or ignoble, if they don't watch their step! Monsters are afoot and the way rife with danger and magic, the path forward may be unclear... But where will is strong, there is a way! Lessons wait in the depths of darkness, and good must prevail. The journey starts now! (Source: FUNimation Entertainment)

First-year high school student Uka Ishimori wants a fresh start. In middle school, Uka was given the nickname Rocky by her peers, being misunderstood as having no emotions when she was actually shy and introverted. As a result, Uka was relentlessly bullied and experienced a lonely, isolated school life. Now, Uka is determined to change. During the first week of school, Uka accidentally gets drenched in lemon soda by her classmate Kai Miura—whose cool personality is the complete opposite of hers. After the incident, Uka is surprised when Kai performs small acts of kindness for her, encouraging her to gradually break out of her shell. Like a lemon soda, bubbly and exciting feelings are beginning to stir.

The aggressive Fuuka Reventon often finds herself in fights as she struggles to make a living. One day, she gets into an altercation with gangsters that leaves her with injuries; luckily, martial arts champion Einhald Stratos finds Fuuka and brings her to a hospital. Seeing potential in Fuuka's skills, Einhald encourages the girl to become a martial arts athlete and offers her a job at Nakajima Gym, where Einhald trains. Despite the kind gesture, Fuuka initially declines the proposal due to the bad memories it brings. She was once childhood friends with the now prominent martial arts athlete Rinne Berlinetta, but the personality of her fellow orphan shifted from kind and gentle to cold and cruel after being adopted by a prosperous family and starting martial arts. Nonetheless, Fuuka soon accepts Einhald's offer and begins training under her—not only out of necessity but also out of the desire to defeat Rinne and make her realize how hurtful her behavior has become. Despite Fuuka's inexperience, Einhald and the other members of the gym have faith in her talent and efforts.

Rumor has it that Momo Kawashima, former student council member at Ooarai Girls Academy, is bound to repeat a year. Having failed her university entry exams, she is far from spending another year at Ooarai, but her plight is unenviable nevertheless. The only way for her to gain admission into a university is to be granted an athletic scholarship. With the Winter Continuous Track Cup rapidly approaching, Momo becomes the commander of Ooarai's tankery team, so as to demonstrate her sport skills and wipe away the stain on her honor. Their first opponent—French-themed BC Freedom High School—is split into two factions that are constantly at odds with each other. Despite this seemingly fortuitous match against a disorganized rival, Ooarai might soon find itself taking one step too far.