Four months before Count von Lohengramm of the Galactic Empire faced Yang Wen-li of the Free Planets Alliance at the Battle of Astarte, he was still just Reinhard von Müsel. The youngest admiral in the Empire's history, Reinhard was disdained and dismissed by his peers as the brother of the Kaiser's concubine. Upon arriving at Iserlohn Fortress with his expeditionary fleet, Reinhard immediately receives an order from the ambitious and cunning Fleet Admiral Gregor von Mückenberger: to intercept an Alliance fleet in a neighboring starzone. Despite recognizing this as a veiled attempt to get him killed in combat, Reinhard nevertheless orders his tired and weary men to engage the enemy in the atmosphere of the gas giant Legnica. But unbeknownst to him, this will mark the first of his many historic encounters with the Hero of El Facil.

In Universal Century 0088, the Anti-Earth Union Group (AEUG) has emerged victorious in its war with the Earth Federation's Titans at the cost of devastating losses. Neo-Zeon, the third faction in the war formerly known as Axis Zeon, remains as powerful as ever. Led by Newtype Haman Karn, Neo-Zeon has been implementing plans to take over both Earth and the space colonies. The AEUG flagship Argama heads to the Side 1 colony Shangri-La for repairs. Living in the colony is Judau Ashta, a 14-year-old junk dealer who is struggling to make enough money to put his younger sister through school. Upon the discovery of an escape pod containing a former Titans pilot, Judau and his friends are quickly led to the Argama in hopes of stealing a mobile suit which they can sell for a fortune. However, with the arrival of a Neo-Zeon ship seeking to defeat the Argama, Judau and his friends are dragged into to a conflict that will bring them across space and Earth.

Following the events of Senki Zesshou Symphogear G, Hibiki Tachibana, Tsubasa Kazanari, and Chris Yukine continue to operate under the command of Genjuurou Kazanari. Meanwhile, Maria Cadenzavna Eve, Kirika Akatsuki, and Shirabe Tsukuyomi are taken into protective custody. With peace restored, everyone returns to their daily lives; however, the appearance of the alchemist Carol Malus Dienheim and her subordinates—the Autoscorers—threatens to draw everyone back into a conflict. Armed with a strange and magical power, Carol wishes to initiate the apocalypse and bring destruction to the world... and she is willing to sacrifice everything to do so. Against this mysterious new foe, the six Symphogear wielders must rise to the challenge in order to protect what they hold dear. However, will Carol and her Autoscorers prove to be too much to handle in this fight to protect the fate of the world?

Deunan, a young female warrior, and Briareos, a veteran cyborg-soldier, are both partners and lovers. As members of E.S.W.A.T., the elite special forces serving Olympus, they are deployed whenever trouble strikes. The two fighters find their partnership tested in a new way by the arrival of a new member to their ranks—an experimental Bioroid named Tereus. (Source: ANN)