Fear, survival, instinct. Thrown into a foreign land with nothing but hazy memories and the knowledge of their name, they can feel only these three emotions resonating deep within their souls. A group of strangers is given no other choice than to accept the only paying job in this game-like world—the role of a soldier in the Reserve Army—and eliminate anything that threatens the peace in their new world, Grimgar. When all of the stronger candidates join together, those left behind must create a party together to survive: Manato, a charismatic leader and priest; Haruhiro, a nervous thief; Yume, a cheerful hunter; Shihoru, a shy mage; Moguzo, a kind warrior; and Ranta, a rowdy dark knight. Despite its resemblance to one, this is no game—there are no redos or respawns; it is kill or be killed. It is now up to this ragtag group of unlikely fighters to survive together in a world where life and death are separated only by a fine line.

An Orb, known only as It, is cast to Earth to be observed from afar. Capable of changing forms from beings whose reflections It captures, It first becomes a rock and then, due to the rising temperature, moss. It does not move until one snowy day, a wolf at death's door barely crosses by. When It takes the animal's form, It attains awareness of its consciousness and starts to wander with an unclear destination in mind. Soon, It comes across the wolf's master—a young boy waiting for his tribe to return from a paradise abundant with fish and fruit in the south. Although the boy is lonely, he still hopes those whom he holds dear in his memories have not forgotten him and that he will reunite with them one day. The boy wants to explore new surroundings and decides to abandon his home with It to find the paradise using the traces his tribe left behind. However, with a heavily injured body and no sight of his elder comrades, what will become of the boy? Fumetsu no Anata e illustrates the story of an immortal being experiencing humanity, meeting all types of people in many places throughout time.

Guts, a man who will one day be known as the Black Swordsman, is a young traveling mercenary characterized by the large greatsword he carries. He accepts jobs that offer the most money, but he never stays with one group for long—until he encounters the Band of the Falcon. Ambushed after completing a job, Guts crushes many of its members in combat. Griffith, The Band of the Falcon's leader and founder, takes an interest in Guts and duels him. While the others are no match for Guts, Griffith defeats him in one blow. Incapacitated and taken into the Band of the Falcon's camp to recover, Guts wakes up two days later. He confronts Griffith, and the two duel yet again, only this time with a condition: Guts will join the Band of the Falcon if he loses. Due to his fresh injuries, Guts loses the fight and is inducted by Griffith. In three years' time, Guts has become one of the Band of the Falcon's commanders. On the battlefield, his combat prowess is second only to Griffith as he takes on large groups of enemies all on his own. With Guts' immense strength and Griffith's leadership, the Band of the Falcon dominate every battle they partake in. But something menacing lurks in the shadows, threatening to change Guts' life forever.

An ancient legend states that with the revival of the Demon God, six heroes—the Braves of the Six Flowers—will be chosen by the Goddess of Fate, granting them power to rise up against the fiends attempting to turn the world into a living hell. Adlet Mayer, self-proclaimed Strongest Man in the World, has arrived at the continent of Piena in hopes of becoming a Brave. Although it doesn't go as smoothly as he had planned, Adlet is ultimately chosen as one of the six heroes shortly after being greeted by Nashetania Loei Piena Augustra, crown princess and fellow Brave. Rokka no Yuusha follows the two as they embark upon their destined journey to fight the Demon God, intending to meet up with their fellow heroes at a small temple outside of the Land of the Howling Demons, the fiends' domain. However, when they finally unite, seven heroes are present, and soon the others begin to suspect Adlet to be a fraud. Now on the run, Adlet must utilize his unique skill set and wit in a fight for his life to identify which member of the group is the true impostor before it's too late!