Following the events of Senki Zesshou Symphogear G, Hibiki Tachibana, Tsubasa Kazanari, and Chris Yukine continue to operate under the command of Genjuurou Kazanari. Meanwhile, Maria Cadenzavna Eve, Kirika Akatsuki, and Shirabe Tsukuyomi are taken into protective custody. With peace restored, everyone returns to their daily lives; however, the appearance of the alchemist Carol Malus Dienheim and her subordinates—the Autoscorers—threatens to draw everyone back into a conflict. Armed with a strange and magical power, Carol wishes to initiate the apocalypse and bring destruction to the world... and she is willing to sacrifice everything to do so. Against this mysterious new foe, the six Symphogear wielders must rise to the challenge in order to protect what they hold dear. However, will Carol and her Autoscorers prove to be too much to handle in this fight to protect the fate of the world?

The story revolves around high school student Akira Nikaido, a typical slacker living a normal life. That is, until he meets the mysterious Shirogane, a man who suddenly appears and tells him that they have a destiny together. When Akira hears this, he is shocked and doesn't believe a word of it. Aya, a friend of Akira, forgets something in the school one night, and asks Akira to help her and go find it. He agrees, and while there, he gets attacked by a shadow monster. Shirogane convinces him that the balance between the human world and the shadow world has been distorted and that Akira must become a shin- a creature of the shadow world- in order to help restore the balance. The anime has shonen-ai themes which are completely absent from the manga. (Source: Wikipedia)

In 2008, the Japanese government began hunting people who identify as otaku in an effort to protect the country's culture. In just one week, the enforcement agency known as the SSC captured almost every otaku by force and confiscated their memorabilia. One passionate man calling himself Otaku Hero refuses to allow this agency to do as they please: he raises a resistance army made up of other displaced geeks who band together to help defend their culture. Three years later, the fires of rebellion wane as all hope seems lost for otakus. Spurred on by his ally—the magical girl Anarchy—Otaku Hero commands the resistance in an effort to reunite Anarchy with the other two other magical girls: Blue and Pink. By combining the power of this trio, the resistance may be able to defeat the SSC and save otaku culture for everyone.

There are rumors that one of Kamome Academy's Seven Mysteries, Hanako-kun, has gained an assistant: first-year high school student Nene Yashiro. Together, Hanako and Yashiro investigate the school's many mysteries, keeping the balance between humans and the supernatural. But in their free time, the two can often be found goofing off, engaging in various antics involving Yashiro's schoolmates and the lurking spectral entities that normal people cannot see.