The fifth and final film in the five-part Gundam: G no Reconguista Movie series.

In Japan, betting on horse racing is one of the few legal forms of gambling; thus, the public races have become an interesting part of Japanese culture. In this sport, an individual who rides a horse and competes in races is called a jockey—that is just what Yuu Arimura aspires to be. Becoming inspired after witnessing a derby, 15-year-old idol Yuu shocks the entire nation when he leaves the entertainment industry to enroll in a prestigious horse racing school. Excited to pursue his latest passion, he gradually bonds with his new classmates—such as the talented Shun Kazanami and the self-assured Amane Grace—who are each unique in their own way. Although the path ahead may be full of obstacles, Yuu is determined to pursue his ultimate goal of becoming one with his horse.

After graduating from high school, Mai Kawai's top priority was to find a job with a stable income to avoid turning out like her father. However, despite having taken a variety of civil service exams, she was faced with a harsh reality: nearly every institute denied her entry. The only place that accepted her was a police academy, which she reluctantly joined. Even now that she is an actual police officer, the world continues to be cruel to Mai. Due to her profession, she unexpectedly receives constant criticism from disgruntled citizens. Disheartened by the hostile treatment, she decides to submit a letter of resignation after just a few months of employment. But on the same day Mai chooses to quit, she meets Seiko Fuji—the former ace of the Criminal Affairs Division who is assigned as her new instructor. Finding her new coworker to be enchanting, Mai delays leaving and gives her uninspiring job a second chance.

Recent college graduate Musashi Shinonome is visiting the Shitamachi God Heaven pachinko hall after inheriting it from his late grandfather. Inside, he stumbles upon a girl sleeping in a futon, who wakes up upon his arrival. She introduces herself as Hanabi Hana Ariake: a self-proclaimed perfected pachislot machine that has turned into a human. To prove this, she demonstrates her ability to transform and shoot fireworks out of her body. Over the following days, Musashi meets two of Ariake's human pachislot machine sisters—the energetic Versus Ikusa Takanawa and the loving Thunder Rai Nihonbashi. Now, Musashi and the dynamic trio must work together to make the grand reopening of Shitamachi God Heaven a success.