After a series of mighty clashes with Upper Rank Demons, the Ubuyashiki clan prepares for one last battle with the hellish forces of Muzan Kibutsuji. In order to finally defeat the Demon leader once and for all, the clan devises a training camp for the Demon Slayer Corps, one led by the remaining Hashira—the most elite warriors in the organization. Each Hashira forms a specialized exercise that will hone both their own abilities and the skills of the ordinary soldiers. Tanjirou Kamado, a boy at the heart of the brewing conflict, recovers from wounds received in a recent fight. While his half-Demon sister Nezuko is studied by researchers like Shinobu Kochou, Tanjirou embarks to train with the Hashira, seeking mastery in each of their assigned areas of expertise to be best prepared for the coming war—skills vital to Tanjirou, as he has vowed to be the very warrior who will eliminate Muzan for good.

For the past 13 years, the Nefarious Monster Army have appeared beneath their floating fortress every Sunday to advance their goal of conquering Earth. Luckily, the Dragon Keepers are here to save the day! Having defeated the majority of the monsters during their initial invasion, the heroes now routinely show up to clear away any remnants of resistance. Unbeknownst to the public, however, every fight beyond the first has been a deliberate show put on by both sides: the Dragon Keepers reap the fame and prestige from fighting imaginary enemies, while the surviving weakest monsters, altogether called Dusters, are allowed to live for another day. Sentouin D, a Duster tired of living this life of shame, leaves the fortress in an attempt to overturn his fate. With the unexpected help of the mysterious ranger Yumeko Suzukiri, he discovers the key to defeating the Dragon Keepers lies in their Divine Artifacts—ultimate weapons crucial to the rangers' transformation and unique powers. Teaming up with Suzukiri to infiltrate the garrison of Red Keeper Sousei Akabane, Sentouin D must defy all expectations to save his species from the cruel hands of humans.

Trapped with thousands of people in The New Gate, a virtual reality game, the elite player Shinya Kiritani resolves to clear the game and finally return home. However, after he manages to single-handedly defeat the final boss, Shinya is transported to a new world—seemingly indistinguishable from the one he just escaped. Shinya discovers that as long as the game menu and his skills are unchanged, he cannot leave this oddly familiar world. In his search of a way to break out from this new dimensional prison, Shinya encounters Tiera Lucent, a hotel receptionist who happens to be a protégée of Schnee Raizar—one of Shinya's former party members. With this glimmer of hope, Shinya strives to reunite with Schnee and find out the truth about The New Gate.

From an early age, Haruka Sakura was made an outcast due to his unconventional appearance and lack of social skills. However, the rough treatment turned him into a proficient fighter, which is now the only thing he prides himself on. Starting at Furin High School, where it is rumored that strength is valued over academics, Sakura has only one goal—taking the top spot. Involved in a street brawl the day before his enrollment, Sakura happens to meet a group of his future schoolmates. Instead of the usual rejection, they fight alongside him, demonstrating that what the school actually cares about is protecting the town of Makochi from any harm—hence why the students call themselves Bofurin. Surprised by the support and appreciation of the townspeople, Sakura has a hard time accepting their goodwill. Though unfamiliar with kindness being shown to him, Sakura must learn to push past his discomfort when Bofurin is pitted against formidable enemies. After experiencing the feeling of acceptance, he finds himself fighting for the sake of others for the first time.