After living peaceful days at the Kamiya dojo, Kenshin Himura finds himself at a crucial crossroads, as his vow to never end a life is put to the ultimate test. Following the murder of his former commander, he now has to retread the road of violence and blood to take down Makoto Shishio—the assassin prodigy who calls himself the notorious Battousai's successor. With ambitions to overthrow the Meiji government and establish his dominion in Kyoto, Shishio has meticulously schemed for a decade, building a formidable anti-government military force. Now, Shishio has turned his sights toward recruiting Aoshi Shinomori, a cold-blooded killer and the former leader of the legendary Oniwaban Group warriors. As Kenshin travels through a village abandoned by the government and currently under the iron fist of Shishio's henchmen, he is determined to prevent the country from spiraling back into civil war. He allies with his former rival, Hajime Saitou, forging a fragile partnership in a battle against Shishio's empire that will determine the fate of Japan.

Aspiring manga artist Mizuho Nishino has grown alongside her four childhood friends: Kizuki Hazawa, Shuugo Hoshikawa, Shin Kashiwagi, and Airu Izumi. But on Mizuho's 17th birthday, Kizuki suddenly asks to be her boyfriend. Having no previous experience with romance, Mizuho brushes it off easily, but Kizuki is genuine and intends to win her heart. Kizuki no longer hides his affections, and Mizuho hesitates, worried about the consequences their potential relationship could have on their whole friend group. As Mizuho works through the confusion of youth, it seems that the other three boys have fallen for her as well—and she must find a way to sort her feelings out without shattering the close-knit group she holds dear.

Tsukasa Akeuraji always dreamed of becoming a competitive solo figure skater, but starting too late in life meant his ambitions never got off the ground. Now barely scraping by, he takes on an assistant coach job, resigned to a future far from the one he once imagined. Before his first day, Tsukasa meets Inori Yuitsuka, a shy fifth grader sneaking into the rink to practice. Captivated by figure skating, but held back by her mother's overprotectiveness and her own self-doubt, Inori has never been encouraged to pursue her passion until she encounters Tsukasa. Beneath her personality lies great potential, waiting to be unlocked with the right guidance. Moved by Inori's determination to change and reminded of his own struggles, Tsukasa offers to coach and help her chase the dream she has been too afraid to voice. While the road ahead is long and filled with unfamiliar challenges and rival skaters, Inori dedicates herself to the art and strives to one day reach the Olympics and become a medalist.

The day Sakiko Togawa's privileged life crumbled, she painfully broke her vow to CRYCHIC and left the band behind—an act that tore her heart in two. As days fade into nights, she struggles to make ends meet until a certain cherished memory is tainted, pushing her to the edge. Driven by a fractured sense of purpose and the need to reclaim what she lost, Sakiko forms Ave Mujica, a masked symphonic metal band. Its members are bound not by a shared goal but by the fragile threads of their divergent desires. Together, they weave a haunting masquerade of loss and rebirth, captivating audiences with a gothic musical stage play. But as the girls' ideals collide, the scars of their pasts rear their ugly heads. The brittle facade the girls have built threatens to collapse beneath them, bringing Sakiko to the verge of falling apart.