It has been 33 years since an alien race, known as the JAM, emerged from a hyperspace passage over Antarctica with the sole purpose of invading Earth. Despite their unexpected initial assaults, a combined effort by humanity not only pushed the JAM force off of Earth but also moved the frontlines of the battle onto the JAM's homeworld of Fairy. Though out of sight and mind for the majority of Earth's inhabitants, the conflict still rages on Fairy, where the United Nations has established five bases and a powerful, technologically advanced military—including the Fairy Air Force (FAF). Lieutenant Rei Fukai acts as a pilot for the FAF Special Air Force’s Boomerang reconnaissance squadron, flying a Super Sylph fighter jet with the personal name Yukikaze. Soaring high above and observing the battles for air superiority, he is responsible for collecting and analyzing valuable intelligence on JAM methods. While his flying skills are unmatched, he also heavily relies on the highly advanced weaponry and near-sentient AI system of his aircraft. Though despised by other members of the FAF for his antisocial and standoffish nature, Lt. Fukai continues to carry out his duties along with Yukikaze. However, during a particularly bloody battle, the appearance of an unresponsive and hostile aircraft catches the lieutenant off-guard—an aircraft which is of the same make and model as his own.

In the cyber city Tom Sawyer, society revolves around earning Love, akin to receiving likes on social media. Citizens of this augmented reality exchange Love for public services and appearance-changing avatars via the Decoration Customizer, also known as Deco, a hologram device implanted in an individual's eyes from early childhood. Under the government's careful management, the people of Tom Sawyer live a peaceful existence, outside the occasional appearance of Phantom Zero, a mysterious criminal who brings all Love to zero within their vicinity. Berry, a mischievous girl fascinated by Phantom Zero, lives comfortably in the system of Tom Sawyer until her Deco malfunctions, enabling her to see a camouflaged prankster named Hack. Believing Hack to be Phantom Zero, Berry chases after the suspect and, in the process, begins to uncover society's most well-kept secrets. As it turns out, Hack is not the only person living outside the system's rules—and the vibrant colors of Tom Sawyer hide a darker nature.

In a world where humanoid armored weapons known as Titatonostrider (TS for short) are used in warfare, troops from various countries converge on the island of Oahu, Hawaii, including Ao Isami of the Japan Ground Self-Defense Force and Lewis Smith of the United States Marine Corps. Isami and Smith cross paths during battle, but suddenly, their teams are attacked by an unknown enemy, scattering soldiers and sending their forces into disarray. In order to save their friends and survive on the deadly battlefield, they must fight with every ounce of courage and pride that they can muster. (Source: Crunchyroll News, edited)

Donning a special mask, a young man defends the planet. (Source: AniDB)