On a fateful day, a young girl named Ai takes a taxi to her new home in Animal Village. After being swindled by her landlord, Tanukichi, into working for his delivery service, it is not long before Ai meets other village residents: Kotobuki, the mayor perpetually worried about his reelection; Yuu, the only other human around, with a passion for random antics and costumes; Bouquet, an overly energetic but rather friendly cat; and Sally, an elephant who dreams of being a fashion designer. New friends seem to lurk in every nook and cranny, but despite that, Ai still struggles to find her niche in this town—until some mysterious notes left on the beach give her a glimpse of a greater purpose.

The friendship between Hiroshi from Japan and Adel from the South Pacific island the two live on is severely tested when Adel finds out that his grandfather was killed in a battle with the Japanese army. Will Adel forget their quarrel when Hiroshi's life is in danger?

Life in Brailsford Junction, Wisconsin, is simple and quiet for Sterling North, who spends most of his free time caring for animals. With his mother in the hospital and his brother fighting on the Western Front, the boy lives only with his father and his dog Hauser. One weekend, cutting away from his usual chores, Sterling, his friend Oscar, and Hauser head off on a fishing trip. On their way to the river, his dog catches a scent at the base of a tree. Unearthing a raccoon's nest, the two boys find an orphaned kit. A passing hunter has killed the kit's mother to make a fur hat, leaving Sterling to adopt and care for the orphaned raccoon. Torn by his fracturing family, a brother in the trenches, along with the ailing health of his mother, Sterling's bonds with life and the world around him are kept together and strengthened by life with his new friend, the ring-tailed wonder and scoundrel, whom he names Rascal.

Aspiring Orange Company Undine Anya Dostoyevskaya feels dejected upon learning that her gondolier mentor, Alice Carroll, is not spending much time with her own mentor, Athena Glory. Athena—the Undine-turned-opera singer—has been attempting to arrange a meeting with Alice, but the latter seems to be avoiding her for unknown reasons. To try and resolve this predicament, Anya calls upon her fellow Undines Ai Aino and Azusa B. McLaren. As the trio works to bring Alice and Athena together, Anya gains a deeper insight into her seniors' feelings, which might just be the key to untangling the uncertainties between them.