Since the night of given's first live performance, the relationship between Mafuyu Satou and Ritsuka Uenoyama has been smooth sailing. Their shared love of music reinforces their bond, and every moment spent together makes them more inseparable. Nevertheless, there will come times when they will suffer setbacks, with both of them consumed by their inner turmoil.

Famous racehorses that have left behind worthy legacies, unique as they can be, are reincarnated as horse girls in a parallel world. In this life, they start their journey anew as they continue to race and perhaps relive the success they once lived through. Aspiring to become the best racehorse in Japan, a horse girl named Special Week moves to Tokyo to enroll in the Tracen Academy—an institution that nurtures horse girls like her to become better racers. There, Special Week witnesses the sophisticated running style of Silence Suzuka and is inspired to become a racer like her. Shortly after, Special Week finds herself recruited into Silence Suzuka's team, Spica. From there, she begins her path to the top—one lap at a time.

After the noble Kappa Kingdom falls to the Otter Empire, the Kappa prince Keppi loses much of his power and becomes helpless against the unseen Kapa-zombies. These zombies plague the world, and are the creations of the Otters and manifestations of people's deepest desires. With no other choice, Keppi must rely on three young boys: Kazuki Yasaka, who must carry a box with him wherever he goes; Enta Jinnai, Kazuki's childhood friend; and Tooi Kuji, a delinquent and a school truant. By having the mythical organ called a shirikodama removed from them, the boys are able to become Kappa themselves and fight the Kapa-zombies. However, to defeat them, the boys must connect with each other via their minds, bodies, and—most importantly—secrets. As the Kappa Kingdom relies on these boys, they must reveal themselves as they have never done before, all the while learning that connections are fragile and truly precious things.

In the warmth of a golden sun, timid high school student Haruka Suzumiya confesses to Takayuki Narumi, the boy she admires. Takayuki accepts Haruka's confession and the two become a couple, supported by their mutual friends Mitsuki Hayase and Shinji Taira. All is well for these friends—until tragedy strikes. One of them is sent to an unenviable fate, lying dormant in a coma. Three years later, they awaken, but their sudden reappearance in everyone's lives threatens to test the limits of these friends' bonds.