Despite not understanding much about fleeting teen romance, first-year high school student Kazuhiko Nukumizu still wonders how he would react if his life were to be turned into a love story. Regardless, as a self-proclaimed background character, Nukumizu is satisfied continuing his life as an introvert with a negligible social life. However, he suddenly finds himself too close to the spotlight when he witnesses his popular classmate Anna Yanami be rejected by her childhood friend in the middle of a family restaurant. While Nukumizu wishes he could just forget what he saw and move on, Anna ends up forcefully confiding herself in Nukumizu, lamenting her status as a childhood friend fated to have her beloved stolen. As he becomes dragged into Anna's situation, Nukumizu soon gets caught up in the relationship drama of two more girls: Lemon Yakishio, an outgoing member of the track and field club; and Chika Komari, a shy member of the literature club. Now thrust out of his comfort zone, Nukumizu finds himself a major character in the lives of too many losing heroines.

Tokiyuki Houjou, a carefree eight-year-old noble, is content with the serene life in Kamakura and shows little regard for the serious responsibilities that come with his eventually succeeding his father as the next shogun. Instead of dedicating himself to rigorous training in swordsmanship or archery, Tokiyuki excels in the art of evasion, skillfully dodging his advisors and discovering perfect hiding spots. However, his peaceful existence is shattered when a sudden coup brutally wipes out his clan. Overwhelmed with guilt for being the sole survivor, Tokiyuki contemplates joining his family in death. However, his fate takes an abrupt turn when the enigmatic priest Yorishige Suwa suddenly shoves him onto a battlefield. Yorishige, who prophesies that Tokiyuki will one day become a great hero, leaves the boy with no choice but to navigate his way free of enemy soldiers. As Tokiyuki struggles to survive against these foes, he finds a new thrill in raising the stakes of his usual hide-and-seek games. With a renewed sense of purpose and the promise of powerful allies from Yorishige, Tokiyuki vows to avenge his family—not through his capability to fight, but through his extraordinary talent for running away.

Much like the other students at her school, first-year student Saki Aoi finds herself captivated by the beauty of her senior on the school board, Makoto Hanaoka. While she worries that Hanaoka would be uncomfortable receiving a confession from another girl, Aoi musters up the courage to tell her anyway. However, Hanaoka's response surprises Aoi for a completely different reason—Hanaoka is actually a cross-dressing boy! Fully expecting Aoi to be abhorred by this revelation, Hanaoka is shocked to find that she has no issues with it. Though Hanaoka rejects Aoi's request to go out with her—citing his identity and the fact that he has never fallen in love before—Aoi insists that she will become his first love. Taken aback by Aoi's determination, Hanaoka wonders if he can really fall in love with her if he cannot find it in him to truly love himself first.

After a series of mighty clashes with Upper Rank Demons, the Ubuyashiki clan prepares for one last battle with the hellish forces of Muzan Kibutsuji. In order to finally defeat the Demon leader once and for all, the clan devises a training camp for the Demon Slayer Corps, one led by the remaining Hashira—the most elite warriors in the organization. Each Hashira forms a specialized exercise that will hone both their own abilities and the skills of the ordinary soldiers. Tanjirou Kamado, a boy at the heart of the brewing conflict, recovers from wounds received in a recent fight. While his half-Demon sister Nezuko is studied by researchers like Shinobu Kochou, Tanjirou embarks to train with the Hashira, seeking mastery in each of their assigned areas of expertise to be best prepared for the coming war—skills vital to Tanjirou, as he has vowed to be the very warrior who will eliminate Muzan for good.