After a series of mighty clashes with Upper Rank Demons, the Ubuyashiki clan prepares for one last battle with the hellish forces of Muzan Kibutsuji. In order to finally defeat the Demon leader once and for all, the clan devises a training camp for the Demon Slayer Corps, one led by the remaining Hashira—the most elite warriors in the organization. Each Hashira forms a specialized exercise that will hone both their own abilities and the skills of the ordinary soldiers. Tanjirou Kamado, a boy at the heart of the brewing conflict, recovers from wounds received in a recent fight. While his half-Demon sister Nezuko is studied by researchers like Shinobu Kochou, Tanjirou embarks to train with the Hashira, seeking mastery in each of their assigned areas of expertise to be best prepared for the coming war—skills vital to Tanjirou, as he has vowed to be the very warrior who will eliminate Muzan for good.

Haruka Kiyomine and Kei Kaname were once recognized as the most formidable pitcher-catcher duo in middle school baseball. Known as the General, Kaname paired his expert strategies with Kiyomine's explosive pitches, crushing the dreams of many starry-eyed players who hoped to one day make it to the big leagues. High school freshman Tarou Yamada is one such former player. Having given up on competing, he enrolls in Kotesashi High School—a completely average public school with no baseball team to speak of—where he discovers that his new classmates are the legendary pair who once destroyed his dreams. Adding to this shocking revelation, he learns that amnesia has reduced the once level-headed General to a vulgar class clown. Having lost all interest in baseball, Kaname dragged a disappointed Kiyomine to Kotesashi simply because the school was close to where they live. As Kotesashi forms a fledgling baseball club, Yamada meets other players he once admired who also left baseball behind. While they might be unwilling now, Yamada believes that if he can convince such talented athletes to start playing again, it might be possible to achieve their dreams and win the national championship—as long as they can put up with Kaname's incessant joking.

Following the faceless god Hitogami's advice seems to have worked wonders for Rudeus Greyrat. After enrolling into the University of Magic as he was told, Rudeus reunites with his childhood friend Sylphiette, who put a valiant effort into curing his condition. The two grow ever closer together and decide to host a wedding party, inviting the friends they have made over the years to announce and formalize their relationship. For all his recent blessings, however, Rudeus' troubles are far from over. The research he is helping Shizuka Nanahoshi conduct hits a bottleneck, sending her into a deep slump much like he experienced in his previous life. Furthermore, a letter from his father, Paul, brings complications to Rudeus' relationships, and Sylphiette still knows next to nothing about his real background. In the face of these issues, Rudeus will have to apply the lessons he has learned in this new world to navigate through the challenges that come with living a life to its fullest.