Shinichi Chiaki conquers his fear of flying, and Megumi Nodame Noda's exceptional performance at a piano competition earns her an invitation to study at the prestigious Conservatoire de Paris. The pair go to Paris together to take the next step in their careers: Chiaki as a new rising conductor under the wing of the great maestro Franz von Stresemann, and Nodame as a pupil of the esteemed piano professor Charles Auclair. But, of course, the music world is much bigger than the two of them could have ever imagined. Chiaki and Nodame, alongside old friends and new rivals, must fight and persevere to reach the dazzling musical heights that await them while never losing sight of what matters most.

Yukihisa Matsuda, principal conductor of the Rising Star Orchestra, arrives in Paris to perform Wagner's Tannhäuser Overture as a guest conductor for the Recyl Symphony Orchestra. When Matsuda overhears the orchestra members praising Shinichi Chiaki, the budding Japanese conductor of the Roux-Marlet Orchestra, he can barely contain his contempt at there being another Black-Haired Noble in the same city. Freshly dumped by his mistress following the performance, Matsuda catches Chiaki and takes him out to drink under the pretense of discussing career. But much to Chiaki's chagrin, the subject soon turns to Chiaki's own special someone.

Shinichi Chiaki is quickly making a name for himself as the principal conductor of the revitalized Roux-Marlet Orchestra, and Megumi Nodame Noda has made leaps and bounds as a pianist at the Conservatoire de Paris. However, tensions mount between the two as Nodame feels left behind by Chiaki's growing success and his close friendship with legendary piano prodigy Rui Son. Disregarding her teacher Professor Charles Auclair's advice, Nodame enters another piano competition in an attempt to jumpstart her own performance career. Meanwhile, those around Chiaki and Nodame are at their own crossroads. Rui begins to doubt herself after hearing Nodame's playing and being denied tutelage from Auclair; Maestro Franz von Stresemann faces the reality of his mortality; pianists Yunlong Li and Tatiana Vishneva feverishly prepare for a competition, while the latter also struggles with her growing feelings for oboist and fellow student Yasunori Kuroki. As Chiaki, Nodame, and their friends continue on their respective journeys, they must not only strive to stay true to themselves, but also remember where it all started.

Short anime specials included in the One Punch Man 2nd Season BD & DVD.